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11. Four early Ordovician continental-shelf assemblages, each indicated by a different symbol. Their positions indicate the presence of biogeographic barriers. After Cocks and Fortey (1990). Fig. 12. Four provinces illustrated by bryozoan distributions: North American (*), Siberian (O), Baltic (O) and Mediterranean ( * ) . Note the puzzling relationships between Baltica (B) and the southeast Asian terranes and between Laurentia (L) and Australia (AU). After Tuckey (1990a). 32 Chapter 2 Unlike the groups so far examined, graptolite biogeography shows that significant changes took place between the early and late Ordovician (Finney and Xu, 1990; Berry and Wilde, 1990).
The former lacks some of the common tropical groups such as calcareous algae, but it is the latter that is the most peculiar. The Malvinokaffric biota contained no stromatoporoids, conodonts, ammonoids, or graptolites and almost no corals or bryozoans. This realm can be recognized until the mid-Devonian when more cosmopolitan conditions began to develop (Boucot, 1988). Later Paleozoic 39 Fig. 15. The three later, early Devonian realms: Eastern Americas (11111),Malvinokaffric (shaded with small open circles) and Old World (shaded with dots).
They somewhat resemble sponges but probably belong in a phylum of their own. These animals, plus calcareous algae, formed waveresistant reefs called bioherms. A little later, in the next stage of the Cambrian, highdiversity shelly faunas, are found. For many years, these remains were thought to represent very small animals. Now, thanks to discoveries in China and Greenland, two larger animals are known that carried the small shells and plates on their bodies (Gould, 1990). The Burgess Shale represents a fortunate accumulation of fine grain sediment in an anoxic environment where scavengers and decay bacteria were absent.